Relay intercropping of subsidiary legumes with durum wheat (living mulch) can be a viable option to support ecological weed control and optimize nutrient cycling in cereal-based cropping systems. However, the lack of knowledge on suitable legume species is often identified as the main bottleneck for the successful application of legume living mulches. This study aimed to evaluate the suitability of 12 different legumes for relay intercropping with wheat in two contrasting Mediterranean cereal-based cropping systems respectively characterized by low-input and integrated management. Each legume was monitored from the undersowing in wheat until the following spring and we compared direct drilling to broadcast sowing of legumes. None of the undersown legumes showed a negative effect on the wheat grain yield. Relay intercropping of legumes proved to be an effective solution to control weeds before and after the wheat harvest, provided suitable legumes species are chosen. Suitable legumes reduced the weed biomass up to the 90% during the intercropping and up to 94% in the following spring. On the contrary, legumes such as Trifoliumresupinatum, Viciavillosa, Medicagotruncatula, and Medicagoscutellata boosted weed growth in the following spring in comparison with the control. According to the performance of legumes, Medicagosativa, Trifoliumrepens and Medicagolupulina had the most suitable characteristics for relay intercropping with durum wheat at the Ravenna site, in a highly productive region whereas Medicagosativa, Hedysarumcoronarium and Trifoliumsubterraneum performed better in the low-input system near Pisa, where yields are generally lower. This is the first time that such a diversity in legumes species is tested in the same experiment for relay intercropping under diversified environmental and management conditions. The results of this study can support farmers in selecting the most appropriated legume species for their specific cropping systems and local conditions.
", "keywords": ["0106 biological sciences", "2. Zero hunger", "Living mulch", "Weed management", "04 agricultural and veterinary sciences", "15. Life on land", "01 natural sciences", "[SDV] Life Sciences [q-bio]", "IWMPRAISE", "Crop combinations and interactions", "Integrated Weed Management", "Horizon2020", "Crop diversification", "0401 agriculture", " forestry", " and fisheries", "Cereals", " pulses and oilseeds", "Subsidiary crops"]}, "links": [{"href": "https://www.iris.sssup.it/bitstream/11382/549112/1/s13593-022-00787-3.pdf"}, {"href": "https://link.springer.com/content/pdf/10.1007/s13593-022-00787-3.pdf"}, {"href": "https://doi.org/10.1007/s13593-022-00787-3"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Agronomy%20for%20Sustainable%20Development", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1007/s13593-022-00787-3", "name": "item", "description": "10.1007/s13593-022-00787-3", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1007/s13593-022-00787-3"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2022-08-01T00:00:00Z"}}, {"id": "10.1016/j.eja.2012.03.006", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:15:25Z", "type": "Journal Article", "created": "2012-04-05", "title": "The Role Of Catch Crops In The Ecological Intensification Of Spring Cereals In Organic Farming Under Nordic Climate", "description": "Abstract Catch crops can contribute to nitrogen supply of following crops through uptake of excess soil mineral nitrogen (N) and through biological N fixation. The contribution of catch crops to the ecological intensification of organic arable systems was investigated using data of a 12-year field experiment carried out at three sites in Denmark. This study focused on the yields of spring oats and spring barley in systems with and without manure in two different cropping systems (O2 and O4) that differed in the proportion of legume-based catch crops (O2 lower and O4 higher) and in the rotation composition (grass\u2013clover green manure in O2 and pulse crops in O4). Three consecutive four-year crop cycles were established at three locations representative of different soil types (loamy sand, sandy loam and coarse sand) and climatic conditions. Crop management and soil operations were performed following common practices in organic farming. Measurements of dry matter (DM) and N content of grain cereals at harvest, above-ground biomass in catch crops and green manure crops in autumn and of the green manure crop at the first cutting were performed. The effect of catch crops on grain yield varied with cereal and catch crop species, soil and rotation type, and the application of N in manure. Higher yield increases from previous catch crops were obtained for spring oat than for spring barley with mean estimates of the apparent N recovery efficiency of N in above-ground catch crops of 69% and 46%, respectively. However, lower autumn N in catch crops undersown in high yielding cash crop was also observed. For spring oats mean grain yield benefits of including catch crops varied from 0.2 to 2.4\u00a0Mg\u00a0DM\u00a0ha\u22121 depending on location, manure use and cycle of the rotation. In spring barley mean grain yield benefits from catch crops varied from 0.1 to 1.5\u00a0Mg\u00a0DM\u00a0ha\u22121. There was a tendency for the effect of catch crop on grain yield to increase over time. These results indicate that in Nordic climates catch crops can contribute to the ecological intensification of spring cereals, not only by reducing the nitrate leaching and increasing N retention, but also by improving yields. Management practices in relation to catch crops must be adapted to the specific soil and cropping systems.", "keywords": ["2. Zero hunger", "Crop combinations and interactions", "legumes", "spring oat", "cropping systems", "15. Life on land", "Cereals", " pulses and oilseeds", "spring barley", "nitrogen"]}, "links": [{"href": "https://doi.org/10.1016/j.eja.2012.03.006"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/European%20Journal%20of%20Agronomy", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1016/j.eja.2012.03.006", "name": "item", "description": "10.1016/j.eja.2012.03.006", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1016/j.eja.2012.03.006"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2013-01-01T00:00:00Z"}}, {"id": "10.1016/j.soilbio.2015.10.008", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:16:12Z", "type": "Journal Article", "created": "2015-11-10", "title": "Evidence For Denitrification As Main Source Of N2o Emission From Residue-Amended Soil", "description": "Catch crops, especially leguminous catch crops, may increase crop nitrogen (N) supply and decrease environmental impacts in cropping systems, but they may also stimulate nitrous oxide (N2O) emissions following spring incorporation. In this 28-day laboratory incubation study, we examined the carbon (C) and N dynamics and N2O evolution after simulated incorporation of residues from three catch crop species into a loamy sand soil, with variable soil moisture (40, 50 or 60% water-filled pore space (WFPS)). The catch crops include two leguminous (red clover and winter vetch) and one non-leguminous species (ryegrass). Plant material was placed in a discrete layer surrounded by soil in which the nitrate (NO3\u2212) pool was enriched with 15N to distinguish N2O derived from denitrification and nitrification. Net N mineralisation from leguminous catch crops was significant (30\u201348\u00a0mg\u00a0N\u00a0kg\u22121 soil, accounting for 41\u201356% of the added residue-N), whereas ryegrass incorporation resulted in net N immobilisation. The evolution of N2O was probably enhanced by N release from the residues, especially during the second week, which can explain the lower N2O evolution after application of ryegrass. Emission of N2O occurred at all moisture levels, but was higher at 50 and 60% WFPS than at 40% in soil with leguminous residues. The 15N enrichment of N2O indicated that denitrification was the dominant source independent of moisture level and residue type. We conclude that catch crop residues will stimulate N2O emissions via denitrification over a wide range of soil moisture conditions, but that emission levels may depend significantly on residue quality and soil moisture.", "keywords": ["Leguminous cover crop", "2. Zero hunger", "Nitrous oxide", "15N labelling", "Nutrient turnover", "Mineralisation", "04 agricultural and veterinary sciences", "incubation", "15. Life on land", "Air and water emissions", "Pasture and forage crops", "Crop combinations and interactions", "13. Climate action", "Farm nutrient management", "Denitrification", "0401 agriculture", " forestry", " and fisheries", "Incubation"]}, "links": [{"href": "https://doi.org/10.1016/j.soilbio.2015.10.008"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Soil%20Biology%20and%20Biochemistry", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1016/j.soilbio.2015.10.008", "name": "item", "description": "10.1016/j.soilbio.2015.10.008", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1016/j.soilbio.2015.10.008"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2016-01-01T00:00:00Z"}}, {"id": "10.1016/j.soilbio.2018.03.008", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:16:13Z", "type": "Journal Article", "created": "2018-03-12", "title": "Forbs Differentially Affect Soil Microbial Community Composition And Functions In Unfertilized Ryegrass-Red Clover Leys", "description": "Increasing plant diversity in agroecosystems is proposed to enhance multiple ecosystem services. Adding selected forbs such as caraway (Carum carvi L.) and plantain (Plantago lanceolata L.) to productive grass-clover mixtures can further enhance forage yields, root biomass, uptake of mineral nutrients and improve animal performance. Yet, it remains unclear whether and how adding these forbs to grass-clover mixtures can influence soil microbial communities and associated soil carbon (C) and nitrogen (N) cycling. Based on a three-year grassland experiment varying in species diversity and composition with and without fertilizer application, we determined soil microbial community composition and functions related to C and N cycling under laboratory incubations. Results showed that inclusion of caraway modified soil microbial community composition by enhancing fungal-to-bacterial phospholipid fatty acids of the ryegrass-red clover mixture. Adding plantain to the ryegrass-red clover mixture increased the relative decomposition rate of the labile C pool, but not of the recalcitrant C pool. Yet, \u03b2-glucosidase activity and net N mineralization were unchanged due to inclusion of either forb. Moreover, fertilization with cattle slurry generally weakened these forb-induced changes in soil microbial properties. These findings demonstrate that adding selected forbs to unfertilized grass-clover leys can modify soil microbial community composition and associated C and N cycling, implying a potential for promoting long-term soil C sequestration through enhanced fungi-to-bacteria ratio, but a limited role in improving soil N fertility.", "keywords": ["2. Zero hunger", "Nitrogen mineralization", "Crop combinations and interactions", "Nutrient turnover", "Fertilization", "Soil carbon dynamics", "Caraway", "0401 agriculture", " forestry", " and fisheries", "04 agricultural and veterinary sciences", "15. Life on land", "Plantain", "Forage mixtures"]}, "links": [{"href": "https://doi.org/10.1016/j.soilbio.2018.03.008"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Soil%20Biology%20and%20Biochemistry", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1016/j.soilbio.2018.03.008", "name": "item", "description": "10.1016/j.soilbio.2018.03.008", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1016/j.soilbio.2018.03.008"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2018-06-01T00:00:00Z"}}, {"id": "10.1079/sum2005326", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:17:04Z", "type": "Journal Article", "created": "2006-03-06", "title": "Pore Characteristics And Hydraulic Properties Of A Sandy Loam Supplied For A Century With Either Animal Manure Or Mineral Fertilizers", "description": "Application of organic residues to soil is generally assumed to improve soil tilth. Only few studies have reported on the long-term effects on the more subtle aspects of soil porosity and no reports have considered the potential effects of organic amendments on the pore system in the subsoil. We sampled undisturbed soil cores (100 cm3 and 6280 cm3) in metal cylinders in differently fertilized plots in the long-term field experiments at Askov Experimental Station, Denmark. We selected the 0-60 cm soil layer of plots dressed for a century with either mineral fertilizers (labelled NPK) or animal manure (labelled AM). Both fertilization treatments were studied at two levels of nutrient application: 'normal' (labelled '1') and 1.5 times 'normal' (labelled '11/2'). Plots unfertilized for a century (labelled UNF) were included as a reference for some of the studies. Water retention, air permeability and air diffusivity were measured on the small cores, and we used the large cores for measuring near-saturated and saturated hydraulic conductivity. In the plough layer, the AM and NPK soils displayed identical pore volumes in size fractions larger as well as smaller than 30 micrometer, while the UNF soil had a significantly smaller volume of pores <30 micrometer. No clear trends were found in treatment effects on pore organization calculated from air diffusivity and air permeability measurements. No significant differences in hydraulic conductivity were found for plough layer soil. For the soil below ploughing depth, significantly larger macropore volumes and near-saturated hydraulic conductivities were found for soil receiving the higher ('11/2') amount of nutrients compared with the 'normally' dressed soil. This effect was independent of fertilization system (AM or NPK). We attribute the larger volume of macropores to the improved root growth conditions in the soil with the larger nutrient level. We conclude that addition of animal manure in rates realistic in agriculture has only a modest effect on soil pore characteristics of the plough layer soil compared with the use of mineral fertilizers. For the soil below ploughing depth, a high level of nutrient application rather than the use of animal manure may increase soil macroporosity and near-saturated hydraulic conductivity.", "keywords": ["2. Zero hunger", "Crop combinations and interactions", "Soil biology", "Nutrient turnover", "Composting and manuring", "0401 agriculture", " forestry", " and fisheries", "04 agricultural and veterinary sciences", "15. Life on land", "Air and water emissions", "Soil quality", "Soil tillage", "6. Clean water"]}, "links": [{"href": "https://doi.org/10.1079/sum2005326"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Soil%20Use%20and%20Management", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1079/sum2005326", "name": "item", "description": "10.1079/sum2005326", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1079/sum2005326"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2005-09-01T00:00:00Z"}}, {"id": "10.1080/00103624.2013.790406", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:17:05Z", "type": "Journal Article", "created": "2013-04-16", "title": "Influence Of Soil Management And Crop Rotation On Physical Properties In A Long-Term Experiment In Parana, Brazil", "description": "This work aims to evaluate the soil physical properties affected by cover crop rotation and soil management in a long-term experiment in southern Brazil. The experiment was established in 1986, with treatments combining six winter treatments and two tillage systems (conventional and no tillage). Bulk density, porosity, aggregate-size class distribution, and organic carbon content of the aggregates were determined at six depths. Bulk density was not affected by tillage systems and winter treatments. The soil disturbance by plowing enhanced the macroporosity, decreased the microporosity, and promoted the formation of smaller aggregate size, in comparison to no tillage. Apart from the soil management, all winter species increased the greater aggregate-size classes, mean weight diameter, geometric mean diameter, and aggregate stability index compared to the fallow treatments. At the no-till treatments, the greater part of sequestered carbon into the soil was stored into the lower and bigger soil aggregates.", "keywords": ["STABILIZATION", "[SDV]Life Sciences [q-bio]", "cover crop", "AGGREGATE", "PARTICULATE", "ORGANIC-MATTER DYNAMICS", "630", "CARBON", "soil organic matter", "Farm nutrient management", "CONSERVATION TILLAGE", "Conservation tillage", "2. Zero hunger", "CULTIVATED SOILS", "04 agricultural and veterinary sciences", "15. Life on land", "sustainability", "Soil tillage", "6. Clean water", "[SDV] Life Sciences [q-bio]", "Crop combinations and interactions", "subtropical soil", "0401 agriculture", " forestry", " and fisheries", "NO-TILLAGE", "CONVENTIONAL-TILLAGE", "FRACTIONS"]}, "links": [{"href": "https://doi.org/10.1080/00103624.2013.790406"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Communications%20in%20Soil%20Science%20and%20Plant%20Analysis", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1080/00103624.2013.790406", "name": "item", "description": "10.1080/00103624.2013.790406", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1080/00103624.2013.790406"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2013-07-20T00:00:00Z"}}, {"id": "10.1111/j.1475-2743.2005.tb00123.x", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:17:41Z", "type": "Journal Article", "created": "2010-08-05", "title": "Nitrate Leaching From Organic Arable Crop Rotations: Effects Of Location, Manure And Catch Crop", "description": "Nitrate leaching from crop rotations supporting organic grain production was investigated from 1997 to 2000 in a field experiment at three locations in Denmark on different soil types. Three experimental factors were included in the experiment in a factorial design: (1) proportion of N2-fixing crops in the rotation (crop rotation), (2) catch crop (with and without), and (3) manure (with and without). Three, four-course rotations were compared, two at each location. The nitrate leaching was measured using ceramic suction cells. Leaching losses from the crop rotation with grass\u2013clover green manure and without catch crops were 104, 54 and 35 kgNha21 yr21 on the coarse sand, the loamy sand, and the sandy loam, respectively. There was no effect of manure application or time of ploughing-in the grass\u2013clover green manure crop on the accumulated nitrate leaching from the entire rotation. Catch crops reduced nitrate leaching significantly, by 30\u201338%, on the sandy soils. At all locations catch crops reduced the annual averaged nitrate concentration to meet drinking water quality standards in the crop rotation with green manure. On the coarse sand there was a time lag between the onset of drainage and the start of N-uptake by the catch crop.", "keywords": ["0106 biological sciences", "2. Zero hunger", "Production systems", "Crop combinations and interactions", "Nutrient turnover", "Composting and manuring", "Farm nutrient management", "0401 agriculture", " forestry", " and fisheries", "Recycling", " balancing and resource management", "04 agricultural and veterinary sciences", "01 natural sciences", "6. Clean water"], "contacts": [{"organization": "M. Askegaard, M. Askegaard, J.E. Olesen, K. Kristensen,", "roles": ["creator"]}]}, "links": [{"href": "https://doi.org/10.1111/j.1475-2743.2005.tb00123.x"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Soil%20Use%20and%20Management", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1111/j.1475-2743.2005.tb00123.x", "name": "item", "description": "10.1111/j.1475-2743.2005.tb00123.x", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1111/j.1475-2743.2005.tb00123.x"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2005-06-01T00:00:00Z"}}, {"id": "10.1111/j.1475-2743.2008.00191.x", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:17:42Z", "type": "Journal Article", "created": "2009-02-25", "title": "Effect Of Different Management Strategies On Soil Quality Of Citrus Orchards In Southern Italy", "description": "AbstractTwenty\uffe2\uff80\uff90six soil samples were collected from 13 paired orchards (organically vs. conventionally managed) homogeneous for age, rootstock and cultivars, belonging to the Eastern Sicily Organic Citrus farm Network. The soil quality was evaluated by chemical and biochemical indicators. The total organic C, humification parameters and isoelectric focusing of extracted organic matter were measured to quantify the size of relevant soil C pools. In addition, C turnover was evaluated by determining microbial C mineralization, C microbial biomass and by calculating the mineralization and metabolic quotient (qCO2). The results obtained demonstrated that organic citrus soils were characterized by a general increase in all the organic matter pools, which means a greater C supply for soil metabolic processes. This observed trend did not directly influence the organic matter turnover, indicating that the organic approach could act as a soil C\uffe2\uff80\uff90sink. The soil microflora of organically managed soils showed an improved efficiency in use of energy and organic resources, corresponding to an increased ability of soils under organic management to sustain biological productivity in the long term.
", "keywords": ["2. Zero hunger", "Crop combinations and interactions", "Nutrient turnover", "Farm nutrient management", "0401 agriculture", " forestry", " and fisheries", "04 agricultural and veterinary sciences", "15. Life on land"]}, "links": [{"href": "https://doi.org/10.1111/j.1475-2743.2008.00191.x"}, {"rel": "related", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/Soil%20Use%20and%20Management", "name": "related record", "description": "related record", "type": "application/json"}, {"rel": "self", "type": "application/geo+json", "title": "10.1111/j.1475-2743.2008.00191.x", "name": "item", "description": "10.1111/j.1475-2743.2008.00191.x", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main/items/10.1111/j.1475-2743.2008.00191.x"}, {"rel": "collection", "type": "application/json", "title": "Collection", "name": "collection", "description": "Collection", "href": "https://repository.soilwise-he.eu/cat/collections/metadata:main"}], "time": {"date": "2009-02-25T00:00:00Z"}}, {"id": "10.2136/sssaj2012.0147", "type": "Feature", "geometry": null, "properties": {"updated": "2026-09-20T16:18:42Z", "type": "Journal Article", "created": "2012-12-19", "description": "In organic cropping systems, legumes, cover crops (CC), residue incorporation, and manure application are used to maintain soil fertility, but the contributions of these management practices to soil nitrogen (N) supply remain obscure. We examined potential sources of N for winter wheat (Triticum aestivumL.) in four experimental cropping systems established in 1997 on three soil types. Three of the four systems were under organic management. Topsoil N, depth of the A horizon, and cumulated inputs of N since 1997 were determined at plot level. Labile soil N pools [mineral N, potentially mineralizable N (PMN), microbial biomass N (MBN)] were monitored during two growth periods; at one site, biomass C/N ratios were also determined. Soil for labile N analysis was shielded from N inputs during spring application to isolate cumulated system effects. Potentially mineralizable N and MBN were correlated across all sites and rotations (r2= 0.72). The MBN corresponded to 46 to 85, 85 to 145, and 74 to 172 kg N ha\uffe2\uff88\uff921at the three sites and differed significantly between cropping systems, but MBN could not explain differences in wheat grain N yields. Instead, a multiple linear regression model explained 76 and 82% of the variation in grain N yields in organic cropping systems in 2007 and 2008, showing significant effects of, respectively, topsoil N, depth of A horizon, cumulated inputs of N, and N applied to winter wheat in manure. Thus, soil properties and past and current management all contributed to winter wheat N supply.
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